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Glossary

abiotic
nonliving
abrade
to wear away through contact
abrasive
tending to wear away through contact
abscission
dropping off or separating
accession
a plant that has been acquired and catalogued
achene
a small, dry, thin-walled type of fruit
actinomycetes
common name for a group of Gram-positive bacteria that are filamentous and superficially similar to fungi
addictive
capable of causing addiction or chemical dependence
adhesion
sticking to the surface of
adventitious
arising from secondary buds, or arising in an unusual position
aeration
the introduction of air
albuminous
gelatinous, or composed of the protein albumin
alkali
chemically basic; the opposite of acidic
alkalinization
increase in basicity or reduction in acidity
alkaloid
bitter secondary plant compound, often used for defense
allele
one form of a gene
allelopathy
harmful action by one plant against another
allopolyploidy
a polyploid organism formed by hybridization between two different species or varieties (allo = other)
alluvial plain
broad area formed by the deposit of river sediment at its outlet
amended soils
soils to which fertilizers or other growth aids have been added
amendment
additive
anaerobic
without oxygen

analgesic
pain-relieving
analog
a structure or thing, especially a chemical, similar to something else
angiosperm
a flowering plant
anomalous
unusual or out of place
anoxic
without oxygen
antenna system
a collection of protein complexes that harvests light energy and converts it to excitation energy that can migrate to a reaction center; the light is absorbed by pigment molecules (e.g., chlorophyll, carotenoids, phycobilin) that are attached to the protein
anthropogenic
human-made; related to or produced by the influence of humans on nature
antibodies
proteins produced to fight infection
antioxidant
a substance that prevents damage from oxygen or other reactive substances
apical meristem
region of dividing cells at the tips of growing plants
apical
at the tip
apomixis
asexual reproduction that may mimic sexual reproduction
appendages
parts that are attached to a central stalk or axis
arable
able to be cultivated for crops
Arcto-Tertiary geoflora
the fossil flora discovered in Arctic areas dating back to the Tertiary period; this group contains magnolias (Magnolia), tulip trees (Liriodendron), maples (Acer), beech (Fagus), black gum (Nyssa), sweet gum (Liquidambar), dawn redwood (Metasequoia), cypress (Taxodium), and many other species
artifacts
pots, tools, or other cultural objects
assayer
one who performs chemical tests to determine the composition of a substance
ATP
adenosine triphosphate, a small, water-soluble molecule that acts as an energy currency in cells
attractant
something that attracts
autotroph
"self-feeder"; any organism that uses sunlight or chemical energy
auxin
a plant hormone
avian
related to birds
axil
the angle or crotch where a leaf stalk meets the stem
axillary bud
the bud that forms in the angle between the stem and leaf
basipetal
toward the base
belladonna
the source of atropine; means "beautiful woman," and is so named because dilated pupils were thought to enhance a woman's beauty

binomial
two-part
biodirected assays
tests that examine some biological property
biodiversity
degree of variety of life
biogeography
the study of the reasons for the geographic distribution of organisms
biomass
the total dry weight of an organism or group of organisms
biosphere
the region of the Earth in which life exists
biosynthesis
creation through biological pathways
biota
the sum total of living organisms in a region of a given size
biotic
involving or related to life
bryologist
someone who studies bryophytes, a division of nonflowering plants
campanulate
bell-shaped
capitulum
the head of a compound flower, such as a dandelion
cardiotonic
changing the contraction properties of the heart
carotenoid
a yellow-colored molecule made by plants
carpels
the innermost whorl of flower parts, including the egg-bearing ovules, plus the style and stigma attached to the ovules
catastrophism
the geologic doctrine that sudden, violent changes mark the geologic history of Earth
cation
positively charged particle
catkin
a flowering structure used for wind pollination
centrifugation
spinning at high speed in a centrifuge to separate components
chitin
a cellulose-like molecule found in the cell wall of many fungi and arthropods
chloroplast
the photosynthetic organelle of plants and algae
circadian
"about a day"; related to a day
circumscription
the definition of the boundaries surrounding an object or an idea
cisterna
a fluid-containing sac or space
clade
a group of organisms composed of an ancestor and all of its descendants
cladode
a modified stem having the appearance and function of a leaf
coalescing roots
roots that grow together
coleoptile
the growing tip of a monocot seedling
collenchyma
one of three cell types in ground tissue

colonize
to inhabit a new area
colony
a group of organisms inhabiting a particular area, especially organisms descended from a common ancestor
commensalism
a symbiotic association in which one organism benefits while the other is unaffected
commodities
goods that are traded, especially agricultural goods
community
a group of organisms of different species living in a region
compaction
compacting of soil, leading to the loss of air spaces
complex hybrid
hybridized plant having more than two parent plants
compound
a substance formed from two or more elements
concentration gradient
a difference in concentration between two areas
continental drift
the movement of continental land masses due to plate tectonics
contractile
capable of contracting
convective uplift
the movement of air upwards due to heating from the sun
coppice growth
the growth of many stems from a single trunk or root, following the removal of the main stem
cortical
relating to the cortex of a plant
covalent
held together by electron-sharing bonds
crassulacean acid metabolism
water-conserving strategy used by several types of plants
crop rotation
alternating crops from year to year in a particular field
cultivation
growth of plants, or turning the soil for growth of crop plants
crystallography
the use of x-rays on crystals to determine molecular structure
cuticle
the waxy outer coating of a leaf or other structure, which provides protection against predators, infection, and water loss
cyanide heap leach gold mining
a technique used to extract gold by treating ore with cyanide
cyanobacteria
photosynthetic prokaryotic bacteria formerly known as blue-green algae
cyanogenic
giving rise to cyanide
cytologist
a scientist who studies cells
cytology
the microscopic study of cells and cell structure
cytosol
the fluid portion of a cell
cytostatic
inhibiting cell division

deductive
reasoning from facts to conclusion
dendrochronologist
a scientist who uses tree rings to determine climate or other features of the past
dermatophytes
fungi that cause skin diseases
desertification
degradation of dry lands, reducing productivity
desiccation
drying out
detritus
material from decaying organisms
diatoms
hard-shelled, single-celled marine organisms; a type of algae
dictyosome
any one of the membranous or vesicular structures making up the Golgi apparatus
dioicous
having male and female sexual parts on different plants
diploid
having two sets of chromosomes, versus having one set (haploid)
dissipate
to reduce by spreading out or scattering
distal
further away from
diurnal
daily, or by day
domestication
the taming of an organism to live with and be of use to humans
dormant
inactive, not growing
drupe
a fruit with a leathery or stone-like seed
dynamical system theory
the mathematical theory of change within a system
ecophysiological
related to how an organism's physiology affects its function in an ecosystem
ecosystem
an ecological community and its environment
elater
an elongated, thickened filament
empirical formula
the simplest whole number ratio of atoms in a compound
emulsifier
a chemical used to suspend oils in water
encroachment
moving in on
endemic
belonging or native to a particular area or country
endophyte
a fungus that lives within a plant
endoplasmic reticulum
the membrane network inside a cell
endosperm
the nutritive tissue in a seed, formed by the fertilization of a diploid egg tissue by a sperm from pollen
endosporic
the formation of a gametophyte inside the spore wall
endosymbiosis
a symbiosis in which one organism lives inside the other

Enlightenment
eighteenth-century philosophical movement stressing rational critique of previously accepted doctrines in all areas of thought
entomologist
a scientist who studies insects
enzyme
a protein that controls a reaction in a cell
ephemeral
short-lived
epicuticle
the waxy outer covering of a plant, produced by the epidermis
epidermis
outer layer of cells
epiphytes
plants that grow on other plants
escarpment
a steep slope or cliff resulting from erosion
ethnobotanist
a scientist who interacts with native peoples to learn more about the plants of a region
ethnobotany
the study of traditional uses of plants within a culture
euglossine bees
a group of bees that pollinate orchids and other rain-forest plants
eukaryotic
a cell with a nucleus (eu means "true" and karyo means "nucleus"); includes protists, plants, animals, and fungi
extrafloral
outside the flower
exudation
the release of a liquid substance; oozing
facultative
capable of but not obligated to
fertigation
application of small amounts of fertilizer while irrigating
filament
a threadlike extension
filamentous
thin and long
flagella
threadlike extension of the cell membrane, used for movement
flavonoids
aromatic compounds occurring in both seeds and young roots and involved in host-pathogen and host-symbiont interactions
florigen
a substance that promotes flowering
floristic
related to plants
follicle
sac or pouch
forbs
broad-leaved, herbaceous plants
free radicals
toxic molecular fragments
frugivous
feeding on fruits
gametangia
structure where gametes are formed
gametophyte
the haploid organism in the life cycle
gel electrophoresis
a technique for separating molecules based on size and electrical charge
genera
plural of genus; a taxonomic level above species

genome
the genetic material of an organism
genotype
the genetic makeup of an organism
germplasm
hereditary material, especially stored seed or other embryonic forms
globose
rounded and swollen; globe-shaped
gradient
difference in concentration between two places
green manure
crop planted to be plowed under to nourish the soil, especially with nitrogen
gymnosperm
a major group of plants that includes the conifers
gynoecium
the female reproductive organs as a whole
gypsipherous
containing the mineral gypsum
hallucinogenic
capable of inducing hallucinations
haploid
having one set of chromosomes, versus having two (diploid)
haustorial
related to a haustorium, or food-absorbing organ
hemiterpene
a half terpene
herbivore
an organism that feeds on plant parts
heterocyclic
a chemical ring structure composed of more than one type of atom, for instance carbon and nitrogen
heterosporous
bearing spores of two types, large megaspores and small microspores
heterostylous
having styles (female flower parts) of different lengths, to aid cross-pollination
heterotroph
an organism that derives its energy from consuming other organisms or their body parts
holistic
including all the parts or factors that relate to an object or idea
homeotic
relating to or being a gene that produces a shift in structural development
homology
a similarity in structure between anatomical parts due to descent from a common ancestor
humus
the organic material in soil formed from decaying organisms
hybrid
a mix of two varieties or species
hybridization
formation of a new individual from parents of different species or varieties
hydrological cycle
the movement of water through the biosphere
hydrophobic
water repellent
hydroponic
growing without soil, in a watery medium
hydroxyl
the chemical group -OH

hyphae
the threadlike body mass of a fungus
illicit
illegal
impede
to slow down or inhibit
inert
incapable of reaction
inflorescence
a group of flowers or arrangement of flowers in a flower head
infrastructure
roads, phone lines, and other utilities that allow commerce
insectivorous
insect-eating
intercalary
inserted; between
interspecific hybridization
hybridization between two species
intertidal
between the lines of high and low tide
intracellular bacteria
bacteria that live inside other cells
intraspecific taxa
levels of classification below the species level
intuiting
using intuition
ionic
present as a charged particle
ions
charged particles
irreversible
unable to be reversed
juxtaposition
contrast brought on by close positioning
lacerate
cut
Lamarckian inheritance
the hypothesis that acquired characteristics can be inherited
lamellae
thin layers or plate-like structure
land-grant university
a state university given land by the federal government on the condition that it offer courses in agriculture
landrace
a variety of a cultivated plant, occurring in a particular region
lateral
to the side of
legume
beans and other members of the Fabaceae family
lignified
composed of lignin, a tough and resistant plant compound
lineage
ancestry; the line of evolutionary descent of an organism
loci
(singular: locus) sites or locations
lodging
falling over while still growing
lytic
breaking apart by the action of enzymes
macromolecule
a large molecule such as a protein, fat, nucleic acid, or carbohydrate
macroscopic
large, visible

medulla
middle part
megaphylls
large leaves having many veins or a highly branched vein system
meiosis
the division of chromosomes in which the resulting cells have half the original number of chromosomes
meristem
the growing tip of a plant
mesic
of medium wetness
microfibrils
microscopic fibers in a cell
micron
one millionth of a meter; also called micrometer
microphylls
small leaves having a single unbranched vein
mitigation
reduction of amount or effect
mitochondria
cell organelles that produce adenosine triphosphate (ATP) to power cell reactions
mitosis
the part of the cell cycle in which chromosomes are separated to give each daughter cell an identical chromosome set
molecular systematics
the analysis of DNA and other molecules to determine evolutionary relationships
monoculture
a large stand of a single crop species
monomer
a single unit of a multi-unit structure
monophyletic
a group that includes an ancestral species and all its descendants
montane
growing in a mountainous region
morphology
shape and form
motile
capable of movement
mucilaginous
sticky or gummy
murein
a peptidoglycan, a molecule made up of sugar derivatives and amino acids
mutualism
a symbiosis between two organisms in which both benefit
mycelium
the vegetative body of a fungus, made up of threadlike hyphae
NADP+
oxidized form of nicotinamide adenine dinucleotide phosphate
NADPH
reduced form of nicotinamide adenine dinucleotide phosphate, a small, water-soluble molecule that acts as a hydrogen carrier in biochemical reactions
nanometer
one billionth of a meter
nectaries
organs in flowers that secrete nectar
negative feedback
a process by which an increase in some variable causes a response that leads to a decrease in that variable

neuromuscular junction
the place on the muscle surface where the muscle receives stimulus from the nervous system
neurotransmitter
a chemical that passes messages between nerve cells
node
branching site on a stem
nomenclature
a naming system
nonmotile
not moving
nonpolar
not directed along the root-shoot axis, or not marked by separation of charge (unlike water and other polar substances)
nonsecretory
not involved in secretion, or the release of materials
Northern Blot
a technique for separating RNA molecules by electrophoresis and then identifying a target fragment with a DNA probe
nucleolar
related to the nucleolus, a distinct region in the nucleus
nurseryman
a worker in a plant nursery
obligate
required, without another option
obligate parasite
a parasite without a free-living stage in the life cycle
odorant
a molecule with an odor
organelle
a membrane-bound structure within a cell
osmosis
the movement of water across a membrane to a region of high solute concentration
oviposition
egg-laying
oxidation
reaction with oxygen, or loss of electrons in a chemical reaction
paleobotany
the study of ancient plants and plant communities
pangenesis
the belief that acquired traits can be inherited by bodily influences on the reproductive cells
panicle
a type of inflorescence (flower cluster) that is loosely packed and irregularly branched
paraphyletic group
a taxonomic group that excludes one or more descendants of a common ancestor
parenchyma
one of three types of cells found in ground tissue
pastoralists
farming people who keep animal flocks
pathogen
disease-causing organism
pedicel
a plant stalk that supports a fruiting or spore-bearing organ
pentamerous
composed of five parts
percolate
to move through, as a fluid through a solid
peribacteroid
a membrane surrounding individual or groups of rhizobia bacteria within the root cells of their host; in such situations the bacteria have frequently undergone some change in surface chemistry and are referred to as bacteroids

pericycle
cell layer between the conducting tissue and the endodermis
permeability
the property of being permeable, or open to the passage of other substances
petiole
the stalk of a leaf, by which it attaches to the stem
pH
a measure of acidity or alkalinity; the pH scale ranges from 0 to 14, with 7 being neutral. Low pH numbers indicate high acidity while high numbers indicate alkalinity
pharmacognosy
the study of drugs derived from natural products
pharmacopeia
a group of medicines
phenology
seasonal or other time-related aspects of an organism's life
pheromone
a chemical released by one organism to influence the behavior of another
photooxidize
to react with oxygen under the influence of sunlight
photoperiod
the period in which an organism is exposed to light or is sensitive to light exposure, causing flowering or other light-sensitive changes
photoprotectant
molecules that protect against damage by sunlight
phylogenetic
related to phylogeny, the evolutionary development of a species
physiology
the biochemical processes carried out by an organism
phytogeographer
a scientist who studies the distribution of plants
pigments
colored molecules
pistil
the female reproductive organ of a flower
plasmodesmata
cell-cell junctions that allow passage of small molecules between cells
polyculture
mixed species
polyhedral
in the form of a polyhedron, a solid whose sides are polygons
polymer
a large molecule made from many similar parts
polynomial
"many-named"; a name composed of several individual parts
polyploidy
having multiple sets of chromosomes
polysaccharide
a linked chain of many sugar molecules
population
a group of organisms of a single species that exist in the same region and interbreed
porosity
openness
positive feedback
a process by which an increase in some variable causes a response that leads to a further increase in that variable

precipitation
rainfall; or the process of a substance separating from a solution
pre-Columbian
before Columbus
precursor
a substance from which another is made
predation
the act of preying upon; consuming for food
primordial
primitive or early
progenitor
parent or ancestor
prokaryotes
single-celled organisms without nuclei, including Eubacteria and Archaea
propagate
to create more of through sexual or asexual reproduction
protist
a usually single-celled organism with a cell nucleus, of the kingdom Protista
protoplasmic
related to the protoplasm, cell material within the cell wall
protoplast
the portion of a cell within the cell wall
psychoactive
causing an effect on the brain
pubescence
covered with short hairs
pyruvic acid
a three-carbon compound that forms an important intermediate in many cellular processes
quadruple hybrid
hybridized plant with four parents
quantitative
numerical, especially as derived from measurement
quid
a wad for chewing
quinone
chemical compound found in plants, often used in making dyes
radii
distance across, especially across a circle (singular = radius)
radioisotopes
radioactive forms of an element
rambling habit
growing without obvious intended direction
reaction center
a protein complex that uses light energy to create a stable charge separation by transferring a single electron energetically uphill from a donor molecule to an acceptor molecule, both of which are located in the reaction center
redox
oxidation and reduction
regurgitant
material brought up from the stomach
Renaissance
a period of artistic and intellectual expansion in Europe from the fourteenth to the sixteenth century
salinization
increase in salt content
samara
a winged seed
saprophytes
plants that feed on decaying parts of other plants

saturated
containing as much dissolved substance as possible
sclerenchyma
one of three cell types in ground tissue
sedimentation
deposit of mud, sand, shell, or other material
semidwarf
a variety that is intermediate in size between dwarf and full-size varieties
senescent
aging or dying
sepals
the outermost whorl of flower parts; usually green and leaf-like, they protect the inner parts of the flower
sequester
to remove from circulation; lock up
serology
the study of serum, the liquid, noncellular portion of blood
seta
a stiff hair or bristle
silage
livestock food produced by fermentation in a silo
siliceous
composed of silica, a mineral
silicified
composed of silicate minerals
soil horizon
distinct layers of soil
solute
a substance dissolved in a solution
Southern blot
a technique for separating DNA fragments by electrophoresis and then identifying a target fragment with a DNA probe
spasticity
abnormal muscle activity caused by damage to the nerve pathways controlling movement
speciation
the creation of new species
specimen
an object or organism under consideration
speciose
marked by many species
sporophyte
the diploid, spore-producing individual in the plant life cycle
sporulate
to produce or release spores
sterile
not capable or involved in reproduction, or unable to support life
sterols
chemicals related to steroid hormones
stolons
underground stems that may sprout and form new individuals
stomata
openings between guard cells on the underside of leaves that allow gas exchange
stratification
layering, or separation in space
stratigraphic geology
the study of rock layers
stratigraphy
the analysis of strata (layered rock)
strobili
cone-like reproductive structures
subalpine
a region less cold or elevated than alpine (mountaintop)

substrate
the physical structure to which an organism attaches, or a molecule acted on by enzymes
succession
the pattern of changes in plant species that occurs after a soil disturbance
succulent
fleshy, moist
suckers
naturally occuring adventitious shoots
suffrutescent
a shrub-like plant with a woody base
sulfate
a negatively charged particle combining sulfur and oxygen
surfaced
smoothed for examination
susceptibility
vulnerability
suture
line of attachment
swidden agriculture
the practice of farming an area until the soil has been depleted and then moving on
symbiont
one member of a symbiotic association
symbiosis
a relationship between organisms of two different species in which at least one benefits
systematists
scientists who study systematics, the classification of species to reflect evolutionary relationships
systemic
spread throughout the plant
tannins
compounds produced by plants that usually serve protective functions, often colored and used for "tanning" and dyeing
taxa
a type of organism, or a level of classification of organisms
tensile forces
forces causing tension, or pulling apart; the opposite of compression
tepal
an undifferentiated sepal or petal
Tertiary period
geologic period from sixty-five to five million years ago
tetraploid
having four sets of chromosomes; a form of polyploidy
thallus
simple, flattened, nonleafy plant body
tilth
soil structure characterized by open air spaces and high water storage capacity due to high levels of organic matter
tonoplast
the membrane of the vacuole
topographic
related to the shape or contours of the land
totipotent
capable of forming entire plants from individual cells
toxin
a poisonous substance
tracheid
a type of xylem cell that conducts water from root to shoot
transcription factors
proteins that bind to a specific DNA sequence called the promoter to regulate the expression of a nearby gene

translocate
to move materials from one region to another
translucent
allowing the passage of light
transmutation
to change from one form to another
transpiration
movement of water from soil to atmosphere through a plant
transverse
across, or side to side
tribe
a group of closely related genera
trophic
related to feeding
turgor pressure
the outward pressure exerted on the cell wall by the fluid within
twining
twisting around while climbing
ultrastructural
the level of structure visible with the electron microscope; very small details of structure
uniformitarian
the geologic doctrine that formative processes on earth have proceeded at the same rate through time since earth's beginning
uplift
raising up of rock layers, a geologic process caused by plate tectonics
urbanization
increase in size or number of cities
vacuole
the large fluid-filled sac that occupies most of the space in a plant cell. Used for storage and maintaining internal pressure
vascular plants
plants with specialized transport cells; plants other than bryophytes
vascular
related to the transport of nutrients, or related to blood vessels
vector
a carrier, usually one that is not affected by the thing carried
vernal
related to the spring season
vesicle
a membrane-bound cell structure with specialized contents
viable
able to live or to function
volatile
easily released as a gas
volatilization
the release of a gaseous substance
water table
the level of water in the soil
whorl
a ring
wort
an old English term for plant; also an intermediate liquid in beer making
xenobiotics
biomolecules from outside the plant, especially molecules that are potentially harmful
xeromorphic
a form adapted for dry conditions
xerophytes
plants adapted for growth in dry areas

zonation
division into zones having different properties
zoospore
a swimming spore
zygote
the egg immediately after it has been fertilized; the one-cell stage of a new individual

Glossary

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